The dependent variable is the rate of the reaction. 

The independent variables are the concentration of enzyme, the concentration of the substrate, the temperature, and the addition of an inhibitor

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R0<1, The population eventually declines to extinction 

R0=1, The population size remains constant

R0>1, The population increases

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λ represents the finite rate of population change over some time interval, often a year

λ = N1/N0

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Populations grow when R0 or λ>1, populations decline when R0 or λ<1, and are at equilibrium when R0 or λ=1

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The simplest method to measure population density is to visually count the number of organisms in a population in a given habitat.

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Quadrats - A square frame often, but not always, measuring 50 × 50 cm.